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A NEW GENUS AND SPECIES OF BLIND CYPRINID FISH FROM CHINA WITH SPECIAL REFERENCE TO ITS RELATIONSHIPS

CHU XINLUO CHEN YINRUI(Kunming Institute of Zoology, Academia Sinica)  
Typhlobarbus, gen. nov.Dorsal and anal without osseous spine. Five branched anal rays. Gill-membranes broadly joined to isthmus. Abdomen rounded. No enlarged scales on both sides of anal base. Two pairs of barbels. Lower pharyngeal teeth in two rows. Air-bladder with two divisions, not enclosed in leathery membrane or bony capusle. Lips simple and smooth, adnate to maxillar and mandible respectively. Eostral fold covering part of upper lip, not fringed at its edge and not forminy a preoral chamber. Rostral groove interrupted at the base of maxillary barbel. Upper lip connected with lower one. Postlabial groove short, only laterally developed and extending to mental groove, which is shallow and inconspicuous. Mental region pad-like, without free rim. Lateral line complete, normally decurved, running in middle of caudal peduncle.Typhlobarbus nudiventris, sp. nov.1. Description The body before dorsal fin is slightly depressed and becomes progressively compressed caudad. Mouth is inferior and arched. Thorax-abdomen is flattened. Eostral barbels reach posterior nostrils and maxillary ones extend as far as the vertical from orbit, which is filled by fat globules except a small opening showing the position of extremely regressed eye. At their symphysis, the dentaries are produced into a small knob, but there is no corresponding notch in the upper jaw. The cephalic lateral line appears to be in two series of pores. Grill rakers are triangle-shaped. Anterior back and thorax-abdomen regions are naked.2. Colouration In life the fish was semitransparent with pale pink color. The branchial region was deep red. The fins were colourless except the tail, which was faint gray with a deeper margin. Abdomen was dark gray, the colour of the stomach content showing through the translucent tissues.3. Locality Three specimens were collected in a dark cavern 100 meters vertically deep below the ground surface at an altitude of 1450 meters. The place is called Yangjieba of Jianshui County, in the south part of Yunnan, China.4. Eelationships Based on morphological comparison with genera Iranocypris, Typhlogarra, Caecocypris and Discogobio the authors come to a conclusion on the systematic position of the new genus Typhlobarbus.1) Typhlobarbus belongs to the subfamily Barbinae of Cyprinidae.2) Though Typhlobarbus may have some affinities with Discogobio in consideration of sharing symplesiomorphies, it is impossible for Typhlobarbus and Discogobio to form a sister group for lack of mutual intercalation between symplesiomorphy and synapomorphy.3) So far the authors have been unable to discover any possible relatives below the subfamily level. This is the reason to erect a new genus to place the new species in.
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