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《Certebrata Palasiatica》 1979-01
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NOTES ON SOME EARLY EOCENE MAMMALIAN FOSSILS OF HENGTUNG,HUNAN

Li Chuan-kuei Chiu Chan-siang Yan De-fa Hsieh Shu-hua (Institute of Vertebrate Paleontology and Paleoanthropology,Academia Sinica)  
A small collection of Early Eocene mammalian fossils was discovered by theauthors in 1976 from three localities(76003,76004 and 76005)in Hengtung County,Hunan.The main fossil quarry(76003)is the very one where C.C.Young collectedremains of Propalaeotherium hengyangensis in 1938.A preliminary observation of boththe old and new materials indicates that:1.Propalaeotherium hengyangensis may be aPropachynolophus rather than a Propalaeotherium;2.the fauna of the upper level(76003and 76004)is certainly Early Eocene in age(Cuisian or even Sparnacian),whereasthe lower one(76005)yielding only two fragments of Archaeolambda mandibles isprobably of the Earliest Eocene;3.the discovery of the two European forms,Pro-pachynolophus and Microparamys in Hunan raises an interesting paleogeographicalproblem:was Eastern Asia isolated perfectly from Europe during the beginning ofEocene,as presumed up to now?1.Propachynolophus hengyangensis(Young);(Equidae,Perissodactyla)Type:a left lower jaw fragment with,M_3(Young's specimen of 1944,V214).Hypodigm:an anterior part of left mandible with P_3—M_2(V 5349).Comparison:The Hunan specimen is similar to Propachynolophus maldani in size,basic cheek teeth structure,degree of premolar molarization and more reduced P_1(cf.Teilhard,1922,p.69),but differs from it by slightly larger size,weaker cingulum,P_3 less molarized and the alveolar margin of the lower jaw anterior to P_8 declinedmore downwards.2.Microparamys lingchaensis sp.nov.(Paramyidae,Rodentia)Type:a lower jaw with a complete tooth row(V5347)Hypodigm:a left lower jaw with M_1—M_3(V 5348)Diagnosis:Size slightly larger than that of M.nanus(P_4—M_3:5.75 mm),masseterfossa end beneath the trigonid of M_3,P_4 small,cheek teeth very primitive,bunodont,with initial ectolophid,anterior cingulum complete,hypoconule small and mesoconiddeveloped.3.Matutinia nitidulus gen.et sp.nov.(Eurymylidae,hnagalida)Type:An anterior part of a brocken skull with complete tooth row(V5354). Hypodigm:five lower jaw fragments(V5359—58,5360),a brocken skull(V5359).Diagnosis:A form intermediate between Heomys and Rhombomylus,premolarsnon-molarized,metaconule rudimentary,posterior margin of the palate ends at the lineof the middle of M~2,paraconid reduced.The enamel of cheek teeth massive.4.Hunanictis inexpectatus gen.et sp.nov.(Didymuconidae,Order indet.)Type:an anterior part of skull with cheek teeth badly crushed(V 5350).Diagnosis:A new genus characterized by its smaller size and more primitivestructure of cheek teeth;protocone of P~2 very small,the inner side of P~4 short,stylesof upper molars appear to be small.5.?Asiocoryphodon sp.(Coryphodontidae,Pantodonta)Only four fragments of lower cheek teeth are preserved(V5351).6.Inseetivora:Represented by two well preserved skulls(V5352-53),detaileddescription of which will be published later.All the measurements of fossils are given in table 1 in the Chinese text.
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